You can trace your 10 coin pay by first noting the wire color going to your payboard for the 10 coin pay. Then, find that same color wire on your reel mech plug, and use your ohm meter to trace the signal back through the reel boards (with single bars on centerline) to the last reel board and probably the grey wire. If it is all working, you can meter it from the pay wire all the way through to the 90 wire on the plug, thus verifying a complete circuit through the reel mech. It sounds like maybe someone changed out one of your reel wipers and forgot to make a notch in the copper foil, or to resolder a notch. These notches btw, separate the boards into different levels, e.g. you don't want 6V bell signal mixing in with a 50V pay signal, so they are separated into isolated circuits by cutting the back of the wipers.
Interesting problem, make certain you keep us informed...