Assuming the payout step up unit is working properly.
The reel wipers may need cleaning and adjusting. Dirty wiper arm assemblies may move slowly or just stay stuck between wiper contacts.
Check wiper arms by manually pulling back then release, they should snap back into the reel index slot.
Testing the reel wipers for continuity.
Start by hooking one side of your meter to the gray wire near top left on the 3rd wiper board.
Run the other lead over the beau plug pins until you get a reading / beep, hook that lead to the pin.
Remove the top lead from the gray wire to confirm the beep should stop. Set up cherries on the pay line.
There should be continuity from the beau plug lead connection across the wipers to the first reel wiper board.
I believe it the white red wire is for the cherries. If no reading try moving the wiper arms slightly back and forth.
This test can be done for all payouts.
Myself once I find continuity for a certain payout I then confirm it at the beau plug.